Research
Interests
Evolutionary Macroecology
Understanding the influences of biotic and
abiotic factors on the evolutionary dynamics of species and clades is
one of
the most
provocative themes in evolutionary biology and paleontology. While
considerable
attention has been
devoted to this subject, the bulk of this work has concentrated on
patterns of
selectivity during mass extinctions and probabilities of turnover in
Recent environments. Relatively little is known about
the
influences of species’ macroecology on evolutionary rates during
the recovery periods following mass extinctions or
those intervals of geologic time characterized by background
intensities of
extinction and
origination. For my doctoral research I am tracking several bivalve
superfamilies (Carditoidea, Pectinoidea, and Veneroidea) from the
Danian through Rupelian
(65 - 30 million years ago) in the Gulf
Coastal Plain
of the southcentral United States
to test causal models of the interactions between species macroecology
and
evolutionary rates during the recovery and diversification of bivalves
following the Cretaceous-Paleogene mass extinction. Whereas previous
studies
have tended to concentrate on individual ecological factors, I am using
path
analysis and structural equation modeling to assess simultaneously the
strength
of direct and indirect interactions between multiple macroecological
characteristics (i.e., abundance, body size, biomass, and geographic
range) and
evolutionary rates as mediated by preservation and sampling. This
research
combines
field
work in the Gulf Coastal Plain, use of museum collections, and data
augmentation from the primary literature.
Abundance and Time-Averaging
The fossil record of faunal abundance may provide the opportunity to
study the ecological and evolutionary responses of
species to environmental change, if the fidelity and acuity of that
record can be determined. Recent meta-analyses of
live-dead data by Kidwell (2001; 2002) have demonstrated that relative
abundance data gathered on skeletal concentrations of marine mollusks
in modern soft-sediment environments can, statistically speaking,
robustly preserve the rank-order, and potentially proportional
abundance, of the live communities they are derived from. In a series
of studies, I have been examining the fidelity of fossilized abundance
through a combination of computer modeling and experimental skeletal
mechanics. Computer modeling can be used to assess the existence,
directionality, and magnitude of post-mortem biases that potentially
affect abundance data in the fossil record. I have used models to
compare the agreement between rank order and eveness in simulated live
and time-averaged death assemblages under a suite of simulated
taphonomic decay regimes. Lab work has focused on the
interplay between generation time, growth rate, and skeletal mechanics
in three species of extant venerid bivalves (Mercenaria mercenaria, Nutricolla tantilla, and Protothaca staminea) to test the
hypothesis that species with short generation times make weaker shells.
Student-Scientist Partnerships
Enabling teachers
and students to participate in the process of doing
science is among the most effective and compelling ways of teaching and
interesting them in science. Participation in actual ongoing scientific
research is perhaps the ultimate example of this kind of hands-on,
inquiry-based involvement. SSPs using local paleontological materials
solve the pedagogical problem of how to get students in touch with real
science
that is interesting to them, connects to their lives and prior
knowledge, but
requires little background or training to make a contribution to
data
acquisition and analysis. While at the Paleontological Research
Institution, my colleagues and I developed the Devonian
Seas Project, an SSP which engaged upper elementary through high
school students and teachers in paleoecology research using the
abundant Devonian marine fossil record of central New York State.
Participants were involved in classroom-based research experiences and
field-based professional development workshops. The
goals of the project included specific substantive educational and
scientific
results and also the creation of a model by which
other
institutions might institute similar partnerships in the geosciences.
To facilitate a dialogue amongst researchers and educators
working
nationally on SSPs, and to aid in the dissemination of project results,
Rob Ross (PRI Director of Education) and I convened two topical
sessions at annual meetings of the Geological Society of America (2001,
2002) and
edited a theme
issue on SSPs in the geosciences for the Journal
of Geoscience Education.
Publications
- Harnik, P.G., in revision.
Unveiling rare diversity by integrating museum, literature, and field
data. Paleobiology.
- Simpson, C., and P.G. Harnik, in review. Assessing the role of
abundance in marine bivalve extinctions over the post-Paleozoic.
Paleobiology.
- Harnik, P.G., and Ross, R.M., 2004. Models of inquiry-based
science
outreach to urban schools. Journal of Geoscience Education 52(5):
420-428. PDF
- Harnik, P.G., and Ross, R.M., 2003. Developing effective
K-16
geoscience research partnerships. Journal of Geoscience Education
51(1): 5-8. PDF
- Harnik, P.G., and Ross, R.M., 2003. Assessing data accuracy
when
involving students in authentic paleontological research. Journal of
Geoscience Education 51(1): 76-84. PDF
- Ross, R.M., Harnik, P.G., Allmon, W.D., Sherpa, J.M.,
Goldman,
A.M.,
Nester, P.L., and Chiment, J.J., 2003. The Mastodon Matrix Project as
an experiment with large-scale collaboration in paleontological
research. Journal of Geoscience Education 51(1): 39-47. PDF
- Simonson, B.M., and Harnik, P., 2000. Have distal impact
ejecta
changed through geologic time? Geology, v. 28, p. 975-978. PDF
|